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XRCC5/XRCC6 polymorphisms in cancers of urinary system

Urinary cancers may include kidney cancer, ureter cancer, bladder cancer, and two

male cancers, testis and prostate cancers. Epidemiologically, all of them seemed to be

more common in male than female. The knowledge about the genomic effects on their

incidence, prognosis, and responses to chemotherapy or radiotherapy is still very

lacking. As for the ureter and testis cancers, there were almost no genomic studies for

the difficulty of sample collection. Followed are the current SNP literatures about

these cancers in urinary system, and the investigations about ureter and testis cancers

are urgently warrant.

3.7.1. Renal cell carcinoma

Renal cell carcinoma is the third leading cause of death among genitourinary

malignancies and the twelfth leading cause of cancer death overall. The incidence of

renal cell carcinoma is about the figure of 2% worldwide. Due to a widespread use of

abdominal imaging, localized tumors are frequently diagnosed nowadays. However,

roughly one third of the patients will ultimately die from this disease even in USA,

where the healthy caring system is very high-qualified.93

In literature, gender, obesity, smoking, analgesic, diuretic abuse, and

environmental factors are reported to be associated with renal cell carcinoma.83

Cigarette smoking, for example, doubles the risk for renal cell carcinoma and

contributes to as much as one third of all cases, yet only a fraction of smokers and a

low number of nonsmokers develop renal cell carcinoma, which implies influence of

host factors on individual susceptibility.94 These individual differences in

susceptibility to renal cell carcinoma may be attributed to genetic polymorphisms in

DNA repair genes or others.95 In 2008, Margulis and his colleagues have investigated

thirteen SNPs in ten DSB repair genes, including XRCC2, XRCC3, NBS1, BRCA2,

RAG1, ATM, and the four NHEJ genes, XRCC5, XRCC6, ligase 4 and XRCC4.96

The original data showed that the SNPs of XRCC5 rs1805388 and XRCC6 rs132788

were not associated with renal cell carcinoma. However, the XRCC6 rs132788

genotype was considered one of the five critical genotypes determining the overall

renal cell carcinoma risk in the classification and regression tree analysis, with the

most effective NBS1 rs1805794. They concluded that individuals carrying more

putative high-risk genotypes in the DSB repair pathway are at higher risks for renal

cell carcinoma.96

3.7.2. Bladder cancer

Bladder cancer is the most common urinary tumor worldwide. In Europe, bladder

cancer is the fourth most frequent cancer among men, accounting for 7% of total

cancers.97, 98 In USA, bladder cancer is the fifth highest cancer in men and seventh in

women.99, 100 Generally, bladder cancer is three times more common in men than

women, and it is primarily a disease of the elderly, with 80% of the patients in the

50-79-year age group and a peak in the seventieth age. Environmental exposures to

tobacco are the predominant risk factors for bladder cancer. The bladder cancer

incidence is two to three folds higher among cigarette smokers as compared with

non-smokers.101 Occupational exposure to carcinogens, alcohol consumption, dietary

factors and the use of hair dyes have also been suggested as risk factors for bladder

cancer.102-106

Although loss of XRCC5 can result in the genome instability and in initialization

of carcinogenesis, over-expression of XRCC5 is associated with the progression of

bladder cancer.107 The expression of XRCC6 is elevated in bladder tumor tissue107 and

XRCC6 may function as a caretaker gene for the development of T-cell

lymphomas.108 In 2008, Wang and his colleagues have published two papers

investigating the roles of XRCC5109 and XRCC6,110 in bladder cancer in China. In the

former study, a polymorphism with a variable number of tandem repeats (21-bp repeat

elements at the position 201 to 160 bp upstream to the initiation of transcription) in

the XRCC5 was investigated of the association with bladder cancer risk. There are

three different alleles, one includes the 42 nucleotide repeat elements (2R), another

contains only one 21-nucleotide repeat in the rectangular box (1R), and still the other

includes no repeat element (0R). The frequencies of the 2R/2R, 2R/1R, and 2R/0R

genotypes among the cases were less than those for the controls, while the proportions

of the 1R/1R, 1R/0R, and 0R/0R genotypes were greater. Overall the difference of the

genotype distributions between the cases and the controls was significant, and

individuals not carrying the 2R allele had a 1.75-fold increased risk of bladder cancer

compared with those carrying the 2R allele.109 The authors has also measured the

promoter activities of the 2R, 1R and 0R alleles by transient transfection in HeLa, T24,

and NIH3T3 cells, finding that fewer tandem repeats in the XRCC5 promoter

increased the activity of the XRCC5 transcript.109 The later paper investigating the

same population has observed an association between XRCC7 rs7003908, but not

XRCC6 rs2267437, genotype and the bladder cancer risk. Also, the risk is increased

among the elder (>65 years old) smokers, suggesting that a gene-environment

interaction may be involved in the development of bladder cancer.110 It is reported that

2-naphthylamine and 4-aminobiphenyl, the compounds in tobacco smoke, can cause

genetic damage in urothelium,111, 112 which may enhance the cellular proliferation in

bladder carcinogenesis.111, 113 In 2009, Michiels and his colleagues have analyzed the

gene-environment joint effects on bladder cancer risk,114 using the classification and

regression tree method similar to that performed gene-gene interaction by Wu.115

Smoking status and genotype data for up to 652 SNPs were incorporated in the

analysis to explore gene-gene and gene-smoking interactions. The outcome is as

expected, the smoking status is the most critical risk factor for bladder cancer. In ever

smokers, a potential two-order interaction between the two SNPs, XRCC5 rs4674066

and ligase 1 rs2288878 was observed. The results suggested that smoking habits,

XRCC5 CC and ligase 1 CT or TT, are sequentially three determinants for bladder

cancer susceptibility in each subject. Very similar to this finding, Chang and his

colleagues has found that there is a joint effects of XRCC5 genotype and personal

smoking habits on bladder cancer risk in Taiwan.116 In this study, a significant

different distribution was found in the frequency of the XRCC5 rs828907, but not

rs11685387 or rs9288518. In addition, those people carried GT and TT genotype at

rs828907 had a 2.05-fold enhanced risk when they had the habit of tobacco smoking,

but not alcohol consumption.116

3.7.3. Prostate cancer

Prostate cancer, a worldwide male disease, is the leading cause of illness and cancer

death in males.117 In addition to age, race and a family history of prostate cancer,

unbalanced diet, androgens, occupational chemicals, smoking, inflammation and

obesity are considered to be additional secondary risk factors.118 Recently, carbon ion

radiotherapy with an established dose fractionation regimen has been shown to yield

biochemically satisfactory relapse-free rates without local recurrence and with

minimal morbidity.119-121 In 2007 and 2008, Suga and his colleagues investigated the

association between 450 SNPs in 118 candidate genes and radiation susceptibility in

prostate cancer patients after carbon ion radiotherapy.122, 123 The genotype of XRCC6

rs2267437, together with those for other four SNPs, SART1 rs2276015, ID3

rs2742946, EPDR1 rs1376264 and PAH rs1226758, were the determinants for the

prediction of developing dysuria after carbon ion radiotherapy in prostate cancer

patients. Despite the small population recruited, their work has set a very good

example for the evaluation of side effects after clinical therapy, using the patients

from a single hospital without the confounding effects of therapeutic protocols and

differential scoring from various examiners and multiple institutions.123

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