56
陸、分類處理
Agrostis L. Sp. Pl. 1: 61-63. 1753.
Annual or perennial. Ligules membranous. Spikelets in panicles, laterally compressed, 1-flowered, articulating above the glumes. Glumes more or less persistent, subequal, membranous, usually 1-nerved, keeled, keel usually scabrous, acute to acuminate. Rachilla process usually absent, or microscopic. Callus glabrous or laterally short-hairy, lemma membranous to scarious, thinner and usually shorter than the glumes, rounded on the back, 5-, rarely 3-nerved, muticous or dorsally awned, apex more or less truncate to 4- or 5-dentate. Palea shorter than the lemma, then
sometimes inconspicuous, veins not or only weakly developed. Lodicules 2, free, thin, veinless, glabrous. Anthers usually 3. Styles 2, free at base; stigmas emerging laterally at anthesis. Caryopsis dorsally grooved to terete; hilum subbasal, punctiform; embryo small.
150~200 species. Worldwide in temperate areas and on tropical mountains.
Key to taxa of Taiwan:
1. Florets with awn
2. Awn 4 mm, geniculated; abaxial of lemma puberulent; anthers 0.3 mm...
...2. A. avenacea 2. Awn 1-2 mm long, straight; abaxial of lemma glabrous; anthers 0.9-1.2 mm
3. Palea 1 mm, 2 nerved; apex of lemma bifid...4. A. dimorpholemma 3. Palea 0.3-0.5 mm; apex of lemma obtuse or truncate
4. apex of palea retuse...7. A. sozanensis 4. apex of palea obtuse...9. A. transmorrisonensis 1. Florets without awn
5. Palea 1 mm, 2 nerved; callus glabrous; anthers 1.2 mm...8. A. stolonifera 5. Palea 0.3-0.8 mm, without nerved
6. Palea 0.3-0.5 mm
7. Anthers 0.3-0.5 mm; callus glabrous...3. A. clavata 7. Anthers 0.6-0.8 mm; callus glabrous or hairy...6. A. infirma 6. Palea 0.6-0.8 mm
8. Anthers 0.6-0.8 mm; callus glabrous, panicle contract, out line lanceolate to oblong...1. A. arisan-montana 8. Anthers 0.9-1.2 mm; callus hairy, panicle erect, outline ovate...
...5. A. fukuyamae
1. Agrostis arisan-montana Ohwi, Acta Phytotax. Geobot. 2: 161. 1933; Hsu, Taiwania 16(2): 229. 1971; Hsu, Taiwan Grasses, 239, pl. 14. 1975; Hsu in Li et al., Fl. Taiwan 5: 398. 1978; Yang in Kuoh, Fl. Reipubl. Popularis Sin. 9(3): 238. 1987;
Lu & Philips in Wu et al., Fl. China 22: 345. 2006. —TYPE: Mt. Arisan, Mar. 7, 1933, J. Ohwi 3463 (Holotype: KYO, n. v.; Isotype: TAI!)
阿里山剪股穎
Agrostis rigidula var. arisan-montana (Ohwi) Veldk., Blumea 28(1): 213. 1982;
Koyama, Grass. Jap. Neighb. Reg. 196. 1987.
Agrostis infirma var. arisan-montana (Ohwi) Veldk., Blumea 41(2): 408. 1996; Kuoh and Chen in Huang et al., Fl. Taiwan, 2nd ed. 5: 341. 2000.
Perennial, tufted. Culms erect, base slender, 30-60cm high, 0.1-2.2 mm in diameter, 4-6 nodes. Leaf-blade flat, linear, 2.1-4.5 mm width, 2.84-13.07 cm long, both surfaces rough, apex acuminate; ligules membranous, apex obtuse, 1.6-3.8 mm long, back scabrous; sheath glabrous. Inflorescence panicle, contracted, lanceolate to narrow oblong in outline, 8.8-21 cm long, 6-7 nodes; branches (2-)8-11(-33) per node, with very short branches and short branchlets, short branches 0.23-1.5 cm, long branches 2.11-8.32 cm, branches rough, semiverticillate, bearing dense spikelets from near the base to apices. Spikelet 1.8-2.2 mm, green or purplish green; glumes
chartaceous, 1-nerveed and keeled, nerve scabrous, apex acute, lower glume slightly longer than the upper glume. Florets 1.35-1.75 mm long; callus glabrous .Lemma membranous, awnless, 5-nerved, apex obtuse or truncate. Palea translucently membranous, 0.62-0.71 mm long. Anthers 0.61-0.73 mm long.
China and Taiwan, on forest margin, roadsides, medium to high altitude.
Specimens examined:
HSINCHU:Leshan-Kuanwu, Jul. 24, 1987, J. C. Wang 4729 (TAI).
TAICHUNG: along the logging trail #710, near Szuyuanyakou, alt. ca. 1900 m, Aug.
15, 2001, W. C. Leong 2398 (HAST); Hsiaohsueh Lodge-Hsiaohsuehshan, alt.
2580 m, C. M. Wang 8385 (TNM); entrance Nanhutashan, Provincial Rd. #9 chia, mileage sign 45 Km, alt. 1933 m, Jun. 2, 2002, I. Liao s.n. (NCKU); Lishan, May 1998, M. I Chen s.n. (NCKU); Szuyuanyakou, Jul, 22, 2008, Y. H. Tsai, 589 (TNU); same loc., Jul. 22, 2008, Y. H. Tsai 592 (TNU).
NANTOU: Yushan, Jul. 10, 1932, S. Suzuki 12987(TAI); Patungkuan-Kuankao, Oct.
16, 1981, T. C. Huang 8507(TAI); Peitungyenshan, Chung Hsing University Horticulture Experimental Farm, alt. 1850 m, Apr. 22, 2002, C. M. Wang 5542 (TNM); intersection of Tatachia and entrance of Chien, Sept. 9, 1986, Y. F. Chen 11388 (NCKU); Chunta logging trail, mileage sign 26 km, Nov. 12, 1986, Y. F.
Chen 12259 (NCKU); Tanta logging trail, mileage sign 25 km, Dec. 1, 2001, K. I.
Liao s.n. (NCKU); Nengkaoyuehling, mileage sign 15-17 km, Oct. 14, 2007, Y. H.
Tsai 382 (TNU).
CHIAYI: Alishan-Tatachia saddle, Oct. 30, 1935, H. Simada 816 (TAI);
Alishan-Chushan, Aug. 12, 1969, C. S. Kuoh 80400(TAI); same loc., Aug. 12, 1969, C. S. Kuoh 80301 (TAI); Alishan, Nov. 16, 1932, Y. Hashioka 142 (TAI);
same loc., Sept. 28, 1967, alt. 2230 m, C. C. Hsu 3903 (TAI); Alishan, Jul. 10, 1933, S. Suzuki 12988 (TAI).
KAOHSIUNG: South Cross Highway, Tienchih, Sept. 20, 1987, C. S. Kuoh 12629 (NCKU); same loc., Jul. 19, 2007, Y. H. Tsai 269 (TNU); intersection of Chinching-bridge to Mt. Kuhanohsin, Aug. 12, 2008, Y. H. Tsai 632 (TNU).
10739 (NCKU); South Cross Highway, Hsiangyang, Jul. 2, 2008, Y. H. Tsai 424 (TNU); same loc., Jul. 2, 2008, Y. H. Tsai 426 (TNU); Mt. Kuanshanling, Jul. 20, 2007, Y. H. Tsai 272 (TNU).
ILAN: Tochiatun, Sept. 3, 2007, Y. H. Tsai 356 (TNU); same loc., Sept. 3, 2007, Y. H.
Tsai 357 (TNU); same loc., Sept. 3, 2007, Y. H. Tsai 370 (TNU).
A. arisan-montana 花序輪廓呈現披針形(Lanceolate)至長橢圓形(Oblong)，花 序分枝具有許多短的分枝和短的次分枝，花序上的小穗密集著生，容易與其他類 群區分。但在檢視標本時發現許多 A. clavata 被錯誤鑑定為 A. arisan-montana，
推測可能原因為Ohwi (1933)原始發表文獻中描述 A. arisan-montana 之花藥長約 1/3 mm，而 A. clavata 花藥長度多介於 0.3~0.5 mm，並且有一型花序具有短分枝，
花序輪廓呈披針形，與 A. arisan-montana 形態非常接近；但根據 Veldkamp (1982)
檢視KYO 模式標本後發現花藥長度約 0.6~0.7 mm，從野外採集及標本館觀察
A. arisan-montana 花藥長度確實不短於 0.6 mm，且內稃長度也明顯可區別此兩 群，A. arisan-montana 內稃 0.6~0.8 mm，A. clavata 內稃 0.3~0.5 mm，詳細比較 見表九。
表九、A. arisan-montana 與 A. clavata 比較
A. arisan-montana A. clavata
花序輪廓 窄披針形至長橢圓形 卵形或窄披針形
花藥長度 0.6~0.7 mm 0.3~0.5 mm
內稃長度 0.6~0.8 mm 0.3~0.5 mm
2. Agrostis avenaceaJ. F. Gmel., Syst. Nat. 1: 171. 1791; Veldk., Blumea 28(1): 207.
1982; M. J. Jung et al., Taiwania 51(1): 25. pl. 1. 2006.
類燕麥剪股穎
Perennial, tufted. Culms erect, to 60-80 cm tall, 3-5-noded, 1.35 mm in diameter.
Leaf-blade flat, linear, about 0.25 mm width, 6.8-10.8 cm long, both surfaces rough, apex acuminate; ligules membranous, apex obtuse, 2.8-4.3 mm long, back scabrous;
sheath glabrous. Panicle inflorescence, effuse, asymmetrical, ovate in outline, 15-25 cm long, 4-5 nodes; branches 3-5 per node, branch 12-18 cm long, scabrous,
verticillate, with spikelets at and near the apices of branches. Spikelet 3-3.25 mm long, green; glumes chartaceous, 1-nerveed and keeled, nerve scabrous, apex acute, lower glume slightly longer than the upper glume. Florets 1.75 mm long. Lemma
membranous, densely hairs all over, puberulent, 5-nerved, apex obtuse, with mid-dorsal awn, geniculated, 4 mm long; callus bearded, hairs 0.4 mm long. Palea translucently membranous, 1.25 mm long, 2-veined, apex bifid. Anthers 0.3 mm long.
Naturalized in southern Taiwan, roadside.
Specimens examined:
KAOHSIUNG: Taoyuan. Teng-chih Forestry Road, Erhchituan, alt. 1400 m, Apr. 19, 2005, M. J. Jung x041903 (NCKU); same loc., alt. 1400 m, Apr. 19, 2005, M. J.
Jung x041906 (NCKU); same loc., alt. 1400 m, Apr. 19, 2005, M. J. Jung x041907 (NCKU); same loc., alt. 1400 m, Apr. 19, 2005, M. J. Jung x041908 (NCKU); same loc., alt. 1400 m, Apr. 19, 2005, M. J. Jung x041904 (HAST);
24, 2008, Y. H. Tasi 406 (NTU); same loc., alt. 1400 m, May 24, 2008, Y. H. Tasi 407 (NTU).
此類群為Jung et al. (2006)發表的歸化植物，目前僅在高雄縣藤枝林道發現，
外稃上被滿毛並具有小穗軸(rachilla)，外稃上具 4 mm 之芒，內稃長 1 mm，具有
兩條脈，先端分叉，花藥0.3 mm，易與其他類群區別。
3. Agrostis clavataTrin. in Spreng., Neue Entd. 1: 55. 1821. Veldk., Blumea 28(1):
209. 1982; Yang in Kuoh, Fl. Reipubl. Popularis Sin. 9(3): 241. pl. 59: 1-4. 1987;
Osada, Illus. Grass. Japan, Enlarged Edition, 304. 1993; Kuoh and Chen in Huang et al., Fl. Taiwan, 2nd ed. 5: 339. pl. 130. 2000.
剪股穎
Agrostis matsumurae Hack. ex Honda, J. Sc. Imp. Univ. Tokyo. 3(1): 191. 1930;
Yang in Kuoh, Fl. Reipubl. Popularis Sin. 9(3): 243. pl. 59 (9-12). 1987 —TYPE:
Japan, Tokyo, Jun. 13, 1880, Matsumura 37 (Holotype: TI, n. v.; Isotype: W).
Agrostis clavata Trin. subsp. matsumurae (Hack. ex Honda) Tateoka, Bull. Nat. Sc.
Mus. Tokyo 11: 161. 1968; Hsu, Taiwania 16(2): 230. 1971; Hsu, Taiwan Grasses, 241. pl. 15. 1975; Hsu, Fl. Taiwan 5: 398. 1978; Osada, Illus. Grass. Japan,
Enlarged Edition, 306. 1993.
Agrostis formosana Ohwi, Bot. Mag. Tokyo 55: 354. 1941. —TYPE: Taiwan, Taihoku, Taiheizan, Mururoahu, Kyanrawa, Mt. Taiheizan, May 26, 1933, J.
Ohwi 2372 (Holotype: KYO, n. v.; Isotype: K).
Perennial, tufted. Culms erect, 22-26 cm tall, 2-5-noded, 0.7-1.6 mm diameter.
Leaf-blade flat, linear, 1.3-3.5 width, 9.6-18.3 cm long, both surfaces rough, apex acuminate; ligules membranous, apex obtuse, 1.4-4.5 mm long, abaxial surface scabrous; sheath glabrous. Panicle either effuse, ovate in outline, or contracted, lanceolate in outline, 7.9-15 cm long, 5-7 nodes; branches 5-6 per node, scabrous, verticillate, long branch 2.08-10.9 cm, short branch 0.4-1.5 cm, with spikelets at and near the apices of branch, shorter ones often spikeled nearly to base. Spikelet
awnless; callus glabrous. Palea translucently membranous, 0.30-0.55 mm long.
Anthers 0.38-0.43(-0.65) mm long.
North Europe, China, Alaska, Japan, Korea and Taiwan.
Specimens examined:
TAOYUAN: mountain top of Papokulushan, alt. ca. 2010 m, Apr. 11, 2002, W. C.
Leong, 2895 (HAST); Lalashan, alt. 1500-2031 m, Apr. 10, 2002, S. C. Wu 2637 (TAI).
HSINCHU: along 100 logging trail, near Yuanyang Lake Nature Reserve, alt. ca.
1685 m, May 18, 2004, C. I Huang 1663 (HAST); Tapachienshan, May 14, 1983, H. I. Chang s.n. (NCKU); Huanchipliou, May 11, 1988, C. S. Kuoh 14397
(NCKU).
MIAOLI:Penglaitsun, Nov. 14, 1972, C. C. Hsu s.n. (TAI); Taian, 230 logging trail mileage sign 12 km, 2500 m, Jul. 27, 1994, C. M. Wang 1124 (TNM); Tunghetsun, alt. 1039 m, Jun. 1, 2002, K. I. Liao s.n. (NCKU); Chialishan, alt. 1420 m, Jun. 1, 2002, K. I. Liao s.n. (NCKU); Ssumahsien logging trail, May 18, 2008, Y. H. Tsai 393 (TNU); same loc., May 18, 2008, Y. H. Tsai 394 (TNU); same loc., May 18, 2008, Y. H. Tsai 395 (TNU); same loc., May 18, 2008, Y. H. Tsai 396 (TNU);
same loc., May 18, 2008, Y. H. Tsai 397 (TNU).
TAICHUNG: Pachsienshan, Aug. 5, 1959, C. C. Hsu 424 (TAI); Ssuyuanyakou, Aug.
22, 1970, C. S. Kuoh 1801(TAI); same loc., Jun. 3, 2007, Y. H. Tsai 168 (TNU);
same loc., Sept. 7, 2007, Y. H. Tsai 371 (TNU); same loc., Jul. 15, 2008, Y. H.
Tsai 618 (TNU); Anmashan, Apr. 20, 1977, C. M. Kuo 8326 (TAI); Changliang Bridge, Provincial Rd. #9 chia, mileage sign 49.2 Km, Jul. 2, 2002, K. I. Liao s.n.
(NCKU); Tahsuehshan logging trail (200 line), mileage sign 34 km, 2031 m, Oct.
19, 2002, C. H. Chen s.n. (NCKU); entrance of Tahsuehshan forest recreation area, mileage sign 35 km, alt. 2016 m, Oct. 20, 2002, C. H. Chen s.n. (NCKU);
Tahsuehshan, Aug. 9, 1986, C. S. Kuoh 10546 (NCKU); Tahsuehshan logging trail, Tenglingchih tunnel, Jul. 26, 2007, Y. H. Tsai 290 (TNU); Tahsuehshan forest recreation area, Tahsuehshan Lodge, Jul. 26, 2007, Y. H. Tsai 291 (TNU);
Provincial Rd. #7 chia, mileage sign 66.4 km, Jun. 2, 2002, K. I. Liao s.n.
(NCKU); Shei-Pa National Park, Hsien Rd. 121, Jun. 3, 2007, Y. H. Tsai 167 (TNU).
NANTOU: Hsitou, Jul. 15, 1954, C. C. Hsu 845 (TAI); same loc., Jul. 15, 1954, C. C.
Hsu s.n. (TAI); Hsitou-Fenghuangshan, Sept. 2, 1969, C. C. Hsu 6098 (TAI);
entrance of Hsitou forest recreation area, Setp. 22, 2001, C. S. Kuoh s.n. (NCKU);
Shanlinhsi, Sept. 23, 1983, S. F. Huang 119 (TAI); Mei-feng, May 16, 1986, M. C.
Tsai 19 (TAI); same loc., Jun. 2, 2007, Y. H. Tsai 169 (TNU); same loc., Jul. 5, 2007, Y. H. Tsai 212 (TNU); Shanlinchi, Apr. 19, 1991, J. G. Liu s.n. (TNM);
Jenai Hsiang: Tunyuan-Yunhai, alt. 1947 m, May 8 2004, Tunghai Univ. Biol.
Coll. Team 9305066 (TNM); Yufeng, Jul. 5, 2007, Y. H. Tsai 214 (TNU);
Nengkaoyuehling, mileage sign 3-4 km, Oct. 11, 2007, Y. H. Tsai 379 (TNU);
Nengkaoyuehling, mileage sign 20-26 km, Oct. 11, 2007, Y. H. Tsai 380 (TNU);
same loc., Oct. 11, 2007, Y. H. Tsai 381 (TNU); en route from Juiyenhsi to Meifeng, Jul. 24, 2008, Y. H. Tsai 596 (TNU); same loc., Jul. 24, 2008, Y. H. Tsai 597 (TNU); Kuanyang, Jul. 24, 2008, Y. H. Tsai 598 (TNU); same loc., Jul. 24, 2008, Y. H. Tsai 599 (TNU).
CHIAYI: Alishan Hsiang, Lichia Chun, en route from Hiking entrance of Toyangshan
1970, C. C. Hsu 8935 (TAI); Alishan, the 3rd branch to the First Branch, Mar. 28, 1970, C. C. Hsu 6863 (TAI); Alishan-Chushan, Aug. 12, 1969, C. S. Kuoh 80390 (TAI); Chaoping, Jul. 12, 1933, Y. Hashioka s.n.(TAI); Alishan, May 17,1932, S.Sasaki s.n. (TAI); same loc., Sept. 1960, C. C. Hsu 1781 (TAI); same loc., Aug.
12, 1969 C. S. Kuoh 80401 (TAI). Shihshan, Feb. 7, 2007, Y. H. Tsai 104 (TNU);
Provincial Rd. #18, mileage sign 91 km, Feb. 7, 2007, Y. H. Tsai 105 (TNU);
Provincial Rd. #18, mileage sign 77 km, Jul. 14, 2007, Y. H. Tsai 255 (TNU);
Tatachia, Jul. 14, 2007, Y. H. Tsai 260 (TNU); same loc., Jul. 14, 2007, Y. H. Tsai 264 (TNU); Fengshan, Jul. 14, 2007, Y. H. Tsai 289 (TNU); Fenchihu, Jun. 2, 2008, Y. H. Tsai 412 (TNU); Fenchihu, Jun. 2, 2008, Y. H. Tsai 413 (TNU);
entrance of Alishan forest recreation area, Jul. 9, 2008, Y. H. Tsai 431 (TNU);
Provincial Rd. #18, Shihtzulu, Jul. 9, 2008, Y. H. Tsai 431 (TNU).
KAOHSIUNG: Shihshan, Jan. 23, 1972, C. C. Hsu 13864 (TAI); Tienchu-Yakou, Jun.
18, 1982, T. C. Huang 8884 (TAI); same loc., Jun. 18, 1982, T. C. Huang 8891 (TAI); Chuyunshan Forest Road, alt. 1000-1100 m, May 7, 2006, C. M. Wang 8915 (TNM); South Cross Highway, mileage sign 124 Km, alt. 1480 m, Apr. 25, 1987, C. S. Kuoh 11556 (NCKU); South Cross Highway, mileage sign 134 Km, Apr. 6, 1988, C. S. Kuoh 14251 (NCKU); South Cross Highway, Tienchi, Dec. 13, 1985, Y. F. Chen 10472 (NCKU); South Cross Highway, Tienchi police station, Jun. 26, 1987, C. S. Kuoh 12306 (NCKU); South Cross Highway, Changchun Bridge, Jul. 19, 2007, Y. H. Tsai 268 (TNU); Tengchih logging trail, May 24, 2008, Y. H. Tsai 408 (TNU); same loc., May 24, 2008, Y. H. Tsai 409 (TNU);
same loc., May 24, 2008, Y. H. Tsai 410 (TNU); same loc., May 24, 2008, Y. H.
Tsai 411 (TNU).
PINGTUNG: Hsiaogueihu logging trail, alt. ca. 1100 m, May 3, 2003, S. M. Ku 1749 (HAST).
TAITUNG: Kuanshanlingshan, Jul. 20, 2007, Y. H. Tsai 268 (TNU).
HUALIEN: Chohsi Hsiang, near Juiwen coal mine, alt. 1300 m, Jun. 22, 1994, C. M.
Wang 1066 (HAST, TNM); Luanshan, Taroko forest, Aug. 16, 1967, alt. 1900 m, C. C. Hsu 3541 (TAI); Hoping Forest road 19.5 km, alt. 1200 m, May 6, 2000, S.
T. Chiu 6145 (TNM); Tayuling, Jul. 23, 2008, Y. H. Tsai 595 (TNU);
Chingshuishan, mountain entrance to Tatung, Jun. 15, 2008, P. F. Lu 14002 (TAIF); Chingshuishan, Chungte, Jun. 15, 2008, P. F. Lu 16072 (TAIF).
ILAN: en route from parking lot to border of Tsuifeng Lodge, alt. ca. 1900 m, Aug. 15, 2001, W. C. Leong 2391 (HAST 90819); on pebbly road between Mt. Fanpaoshan and Shenmihu, alt. ca. 1390 m, Aug. 14, 2003, M. W. Jian 112 (HAST);
Chilanshan logging trail, at road mileage sign 22 km, alt. ca. 1950 m, May 2, 2002, C. H. Lin 197 (HAST, TNM); Tatung, Chialohu, alt. 2230m, May 12, 2002, C. I. Huang 886 (HAST); Taipingshan, Apr. 25, 1930, S. Sasaki s.n. (TAI); same loc., Jun. 29, 2007, Y. H. Tsai 223 (TNU); same loc., Jun. 29, 2007, Y. H. Tsai 224
Chililoshan, Jul. 16, 1932, T. Suzuki 7304 (TAI); same loc., May 12, 1932,
S.Sasaki s.n. (TAI); Tatung, 101 forest road 1-2 K, alt. 1100 m, Jul. 15, 2003, C. L Huang 164 (NTM); Tatung, 100 forest road 12-13K, Divine tree Area, alt. 1600 m, Jul. 15, 2003, C. L Huang 147 (TNM); same loc., Jul. 15, 2003, C. L Huang
A. clavata 過去主要常因花序變化將之區分為兩分類群，其一具鬆散圓錐花 序，小穗著生在分枝頂端或上半部，分枝下半部裸露無小穗，為 A. clavata；另 一群為緊縮圓錐花序，具短分枝，小穗著生在分枝和次分枝的基部至先端，分枝 無裸露，具有此花序特徵多被提出為 A. clavata 種下分類群或是提升為種(A.
matsumurae)。但兩類型除花序形態外並無其他可區別特徵，而且曾經觀察在同 一叢植株就存在兩種花序類型(圖四 C)，因此 A. clavata 雖然具有花序上變化，
但不足以做為種甚至種下分類差別。
A. clavata 為廣泛分布類群，西伯利亞、北美洲阿拉斯加、中國大陸、日本皆
有分布，台灣地區北部低海拔至中南部海拔3000 m 以下很容易見到此類群。
4. Agrostis dimorpholemma Ohwi, Bot. Mag. Tokyo 55: 351. 1941; Koyama, Grass.
Jap. Neighb. Reg. 197. 1987; Osada, Illus. Grass. Japan, Enlarged Edition, 310. 1993;
Kuoh, Taiwania 48(2): 94. pl. 1. 2003. —TYPE: Japan, Hondo, Hakono-Asinoyu in Sagami, Jul. 1930, K. Hisauchi s.n. (Holotype: KYO, n. v.)
多形剪股穎
Perennial, with slender stolons. Culms erect from a shortly prostrate or geniculate ascending base, 50-80 cm tall, 1.1-2.5 mm diameter, with 4-6-nodes. Leaf-blade flat, linear, 1.5-3.2 mm width, 11.3-13.1 cm long, both surfaces glabrous, apex acuminate;
ligules membranous, apex obtuse, 1.35-3.92 mm long, back scabrous; sheath glabrous.
Panicle effuse, ovate in outline, 16-25 cm long, 4-5 nodes; branches 3-5 per node, branch 2.4-12 cm long, branch scabrous, verticillate, with spikelets at and near the apices of branch, and lower 1/2-1/3 branch bare. Spikelet 2.4-2.6 mm long, green and often tinged with purple. Glumes chartaceous, 1-nerve and keeled, nerve scabrous, apex acute, lower glume slightly longer than the upper glume. Florets 1.8-2.2 mm long, dimorphic, with awned and awnless ones on the same branch; awnless lemma membranous, 3-nerved, sometimes midribs with a tooth arising from near the apex or middle of the lemma, glabrous, apex 2-toothed, callus glabrous; awned lemma membranous, 3-nerved, apex 2-toothed, awn arising from near the base, awn 1.5-2 mm long, long hair on the basal part and on callus. Palea translucently membranous, 0.98-1.2 mm long, apex bifid, 2-veined. Anthers 0.9-1.25 mm long.
Distributies in Japan and Taiwan.
Specimens examined:
x062713 (NCKU). Provincial Rd. #18 Mileage sign 95 km, Jul. 14, 2007, Y. H.
Tsai 258 (TNU); Provincial Rd. #18, Shihshan Station, Jul. 14, 2007, Y. H. Tsai 261 (TNU). Provincial Rd. #18, entrance Alishan Forest Recreation Aera, Jul. 9, 2008, Y. H. Tsai 429 (TNU).
Agrostis dimorpholemma 為 Kuoh (2003)發表台灣的新紀錄種，目前僅在嘉義
阿里山往南投信義一帶有分布。在同一花序上具兩型小花，外稃有芒，芒1 mm
長，基盤有毛，另一型為外稃無芒或具短突的，基盤通常光滑，外稃先端凹呈二 齒形，花藥1 mm，內稃 1 mm 長。Agrostis dimorpholemma 一直被認為是天然雜 交類群(Ohwi, 1941; Koyama, 1987; Osada, 1993)，但對於雜交的親本仍是未知，
在形態觀察由於具有匍匐莖，應可以藉由營養繁殖傳播、繁衍。
5. Agrostis fukuyamae Ohwi, Fedde. Repert. 36: 39. 1934; Hsu, Taiwania 16(2): 230.
1971; Hsu, Taiwan Grasses, 243. pl. 15. 1975; Hsu, Fl. Taiwan 5: 400. pl. 1376. 1978;
Yang in Kuoh, Fl. Reipubl. Popularis Sin. 9(3): 240. 1987; Lu et al. in Wu et al., Fl.
China 22: 345. 2006. —TYPE: Mt. Nanhutashan, Jul. 19-20, 1933, J. Ohwi 4147 (Holotype: KYO, n. v., Isotype: K, n. v.); same loc., N. Fukuyama 4090 (Paratype:
TAI!).
伯明剪股穎
Agrostis rigidula var. fukuyamae (Ohwi) Veldk., Blumea. 28(1): 219. 1982; Koyama, Grass. Jap. Neighb. Reg. 197. 1987.
Agrostis infirma var. fukuyamae (Ohwi) Veldk., Blumea. 41(2): 408. 1996; Kuoh and Chen in Huang et al., Fl. Taiwan, 2nd ed. 5:343. 2000.
Perennial, tufted. Culms erect, 11-24 cm tall, 2-4-noded, 0.75-0.94 mm diameter.
Leaf-blade flat or involute, linear, 0.55(involute)-2.07 width, 4.78-13.03 cm long, both surfaces rough, apex acuminate; ligules membranous, apex obtuse, 1.52-4.08 mm long, back scabrous; sheath glabrous. Panicle effuse, ovate in outline, 7.38-20 cm long, 5-7 nodes; branches 4-7 per node, branch 2.7-8 cm, scabrous, verticillate, with spikelets at and near the apices of branch. Spikelet 1.9-2.5 mm, purplish green;
glumes chartaceous, 1-nerveed and keeled, nerve scabrous, apex acute, lower glume slightly longer than the upper glume. Florets 1.47-1.85 mm long. Lemma
membranous, glabrous, 5-nerved, apex obtuse or truncate, awnless; callus minutely hairy. Palea translucently membranous, 0.72-0.75 mm long. Anthers 0.91-1.18 mm long.
Endemic to Taiwan at high altitude.
HSINCHU:Itseshan, Jul. 18, 1932, S.Sasaki s.n. (TAI); Tapachienshan, alt.
2700-3500 m, Aug. 2, 1972, C. S. Kuoh 3266 (TAI).
TAICHUNG: Nanhutashan, Taosaifeng, alt. 3400 m, Jul. 15, 2002, W. C. Leong 3216 (HAST); Nanhutashan, Jul. 1933, N. Fukuyamae 4090 (TAI); Nanhutashan, Sept.
5, 2007, ; Tapachienshan, Aug. 2, 1972, C. C. Hsu 12143 (TAI); 369 shanchuang to Hsuehshan, Oct. 24, 1936, K. Mori s.n. (TAI); Hsuehshan, Jul. 23, 1932, S.
Sasaki s.n. (TAI); same loc., Aug. 4, 2007, Y. H. Tsai 282 (TNU); same loc., Aug.
4, 2007, Y. H. Tsai 283 (TNU); same loc., Aug. 17, 2008, Y. H. Tsai 626 (TNU);
Nanhuchufeng, Sept. 5, 2007, Y. H. Tsai 360 (TNU); Kupo to Hsuehshantungfeng, Jul. 16, 2008, Y. H. Tsai 473 (TNU); Kupo to Hsuehshantungfeng, Jul. 16, 2008, Y.
H. Tsai 480 (TNU); Hsuehshantungfeng to 369 shanchuang, Jul. 16, 2008, Y. H.
Tsai 490 (TNU); same loc., Jul. 16, 2008, Y. H. Tsai 491 (TNU); same loc., Jul. 16, 2008, Y. H. Tsai 492 (TNU); same loc., Jul. 16, 2008, Y. H. Tsai 496 (TNU); same loc., Jul. 16, 2008, Y. H. Tsai 497 (TNU); same loc., Jul. 16, 2008, Y. H. Tsai 504 (TNU); Wuyenfeng to Yunleng Lodge, Sept. 6, 2007, Y. H. Tsai 367 (TNU).
NANTOU:Chilaichushannanfeng, Aug. 24, 1929, S.Sasaki s.n. (TAI); same loc., Aug.
24, 1929, S.Suzuki 2229 (TAI); same loc., Aug. 24, 1929, S.Suzuki 2300
(TAI);same loc., Aug. 24, 1929, S.Suzuki 2304 (TAI) ;same loc., Aug. 24, 1929, S.Sasaki 380176 (TAI); Nengkao, Aug. 4, 1926, S.Suzuki s.n. (TAI); same loc. Jul.
14, 1930, Y. Kudo 126; Kuailin-Nengkao, Aug. 23, 1929, S. Sasaki s.n. (TAI);
Wuling, Jul. 6, 2007, Y. H. Tsai 217 (TNU); same loc., Jul. 6, 2007, Y. H. Tsai 219 (TNU); same loc., Jul. 24, 2008, Y. H. Tsai 611 (TNU); same loc., Jul. 24, 2008, Y.
H. Tsai 614 (TNU); mountain entrance of Hohuantungfeng, Jul. 6, 2007, Y. H.
Tsai 218 (TNU).
KAOHSIUNG:South Cross Highway, Hsiangyang, Jun. 24, 1988, C. S. Kuoh 12039B (NCKU).
A. fukuyamae 為台灣特有種，目前其他地區尚未記錄此類群分布，主要分布
地點在海拔3000 m 以上，生育環境為開闊且陽光充足的草坡、步道兩旁或是高
海拔岩屑基質，常與 A. infirma 和 A. sozanensis 共域分布。花藥約 1 mm，內稃約 0.8 mm，基盤有毛等特徵容易與其他類群區別。
Hsu (1971, 1975, 1978)皆有記錄 A. fukuyamae，但是其檢索表指出 A.
fukuyamae 外稃有芒，而內文描述為無芒；而依據 Ohwi (1934)原始發表文獻，以 及檢視TAI 所藏的一份複模式(Paratype)，皆顯示 A. fukuyamae 外稃不具芒，因 此推測Hsu 的檢索表所指的類群，可能包含了內稃較明顯的 A. sozanensis，也符 合許多 A. sozanensis 標本被錯誤鑑定為 A. fukuyamae 的情況，應是使用 Hsu 檢索 表所致。
6. Agrostis infirma Buse in Miq., Pl. Jungh. 3: 342. 1854; Veldk., Blumea. 41(2): 408.
1996; Lu and Phillips in Wu et al., Fl. China 22: 345. 2006. —TYPE: Indonesia, Java, F. W. Junghuhn s.n. (Holotype: L-903.342-37, n. v.)
玉山剪股穎
Agrostis flaccida Hack. var. morrisonensis (Hayata) Honda, Bot. Mag. Tokyo 40: 324.
1926.
Agrostis morrisonensis Hayata, Icon. Pl. Form. 7: 86. pl. 53. 1918; Hsu, Taiwania 16(2): 230. 1971; Hsu, Taiwan Grasses, 245. 1975; Hsu, Fl. Taiwan 5: 400. 1978;
Yang in Kuoh, Fl. Reipubl. Popularis Sin. 9(3): 243. pl. 59: 13-16. 1987. —TYPE:
T. Kawakami et U Mori 2374 (Syntype: not traced); Nov. 1906, T. Kawakami et U Mori 2365 (Syntype: not traced).
Agrostis rigidula Steudel, Synops. Pl. Glumac. 1: 171. 1854; Veldkamp, Blumea.
28(1): 219. 1982; Koyama, Grass. Jap. Neighb. Reg. 196. 1987. —TYPE:
Indonesia, Java, Zollinger 2589 (Holotype: P).
A. clarkei auct. non Hook.: Hayata, Fl. Mon. Form. 237. 1908.
Perennial, tufted. Culms erect, 10.3-43 cm tall, 3-4-noded, 0.94-1.27 mm diameter. Leaf-blade flat or involute, linear, 1.48-2.08 mm width, 2.69-10.7 cm long, both surfaces rough, apex acuminate; ligules membranous, apex obtuse, 1.55-2.87 mm long, back scabrous; sheath glabrous. Panicle effuse, ovate in outline, 7.9-15 cm long, 5-7 nodes; branches 5-6 per node, branch 2.08-10.9 cm, scabrous, verticillate, with spikelets at and near the apices of branch. Spikelet 1.5-2.6 mm, purplish green;
glumes chartaceous, 1-nerved and keeled, nerve scabrous, apex acute, lower glume
glumes chartaceous, 1-nerved and keeled, nerve scabrous, apex acute, lower glume