A List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) Deposited in the Insect Collection of Taiwan Agricultural Research Institute

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(1)台灣農業研究 (J. Taiwan Agric. Res.) 63(2):105–121 (2014) DOI: 10.6156/JTAR/2014.06302.02. A List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) Deposited in the Insect Collection of Taiwan Agricultural Research Institute Shu-Pei Chen1,*, Jen-Yu Wong2, and Wen-Jer Wu3,4. Abstract Chen, S. P., J. Y. Wong, and W. J. Wu. 2014. A list of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) deposited in the Insect Collection of Taiwan Agricultural Research Institute. J. Taiwan Agric. Res. 63(2):105–121.. A list of 39 species in 9 genera of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) with 246 slide mounted specimens deposited at Insect Collection of the Taiwan Agricultural Research Institute (TARI) is provided. The specimens were mainly collected during Japanese Colonial Period. The synonyms, host plants, distribution and photographs of slide-mounted specimen are also provided for each species. Key words: Chionaspinae (Diaspididae), Slide-mounted specimens, Taiwan Agricultural Research Institute (TARI).. INTRODUCTION Scale insects (Hemiptera: Sternorrhyncha: Coccoidea) of Taiwan have been extensively studied by serveral western and Japanese entomologists such as Green, Ferris and Takahashi since 1923 (Tao 1978). Most of the slidemounted specimens of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) collected by Takahashi were deposited at the Insect Collection, Division of Applied Zoology, Taiwan Agricultural Research Institute (TARI). Among them, a great number of slidemounted specimens were types designated by Takahashi. However, a considerable number of species names used by Takahashi have been chang ed or synonymized with other names.. In this study, taxonomic status of all slidemounted specimens were examined and revised based on the current studies.. MATERIALS AND METHODS A total of 246 slide-mounted specimens were examined by using a Leita DMRB light microscope (Leica, Milton Keynes, U.K.) in this study. Distribution data of each species were compiled based on the information provided on the slide-mounted specimens deposited at the TARI Insect Collection. Photographs of slidemounted specimens (adult female) were taken by using Leita DMRB light microscope, Automatic Piezo stepping motor (Leica Piezo Zdrive, Switzerland) and Leica DFC 420 Digital. Received: January 24, 2014; Accepted: March 26, 2014. * Corresponding author, e-mail: spchen@tari.gov.tw 1 Associate Research Fellow, Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung, Taiwan, ROC. 2 Former Field Technician, Applied Zoology Division, Taiwan Agricultural Research Institute, Taichung, Taiwan, ROC. 3 Professor, Department of Entomology, National Taiwan University, Taipei, Taiwan, ROC. 4 Director, Research Center for Plant Medicine, National Taiwan University, Taipei, Taiwan, ROC..

(2) 106. 台灣農業研究第 63 卷第 2 期. Microscope Cameras (Leica, Switzerland) for each species. The old name and current name of collection locality records are included in the paper based on guidelines developed by Chu & Yamanaka (1973, 1974, 1975) and the Taiwan Geographic Name Information Systems (http:// placesearch.moi.gov.tw/search/index.php).. RESULTS Among the identified 39 species (9 genera) of Chionaspinae in TARI, there were 22 species described by Takahashi, 5 species described by Kuwana, and 5 species described by Green. In the study, there are 10 species not collected in Taiwan, including Chionaspis lithocarpicola Takahashi, Chionaspis salicis (Linnaeus), Kuwanaspis linearis (Green), Nikkoaspis sasae (Takahashi), Nikkoaspis shiranensis Kuwana, Pinnaspis javanensis (Kuwana), Pinnaspis siamensis Takahashi, Unaspis acuminata (Green), Unaspis citri (Comstock), and Unaspis euconymi (Comstock). The following list provides comprehensive information on the nomenclature, synonymies, host plants, and distribution of each species. Table 1 is a check list of the old name and current name of collection locality records included in the paper (Chu & Yamanaka 1973, 1974, 1975). Chionaspis cinnamomicola (Takahashi 1935) (Fig. 1A) (Takahashi 1935a; Takagi 1970; Tao 1989; Ben-Dov & Miller 2013). Original spelling on label: Diaspis machilicola var. cinnamomicola Takahashi, 1935. (8 slides) Distribution: Taiwan (Giran: Kussha; Takao: Chushinron near Rokki). Host plant: Cinnamomum erythrophloia (Lauraceae). S y n . : D ia s p is m a ch ilico la cin n a m o m ico la Takahashi, 1935; Diaspis cinnamomicola (Takahashi 1935).. Note: Syntypes, female [six slides collected from Taiwan: Taihoku, Kussha near Shikikun, on Cinnamomum erythrophloia, 14-VIII-1934, by R. Takahashi ( #26-3-31–26-3-4-1)]. Chionaspis lithocarpicola Takahashi, 1942 (Fig. 1B) (Takahashi 1942b; Ben-Dov & Miller 2013) Original spelling on label: Chionaspis lithocarpicola Takahashi, 1942 (1 slide) Distribution: Thailand (Mt. Sutep). Host plant: Lithocarpus sp. (Fagaceae). Note: Syntypes, female [collected from Thailand: Mt Sutep, on Lithocarpus sp., 12IV-1940, by R. Takahashi ( #23-4-4-1)]. Chionaspis machilicola (Takahashi 1935) (Fig. 1C) (Takahashi 1935a; Takagi 1970; Tao 1989; Ben-Dov & Miller 2013). Original spelling on label: Diaspis machilicola Takahashi, 1935 (4 slides) Distribution: Taiwan (Taito: Chipponsan). Host plant: Machilus sp. (Lauraceae). Syn.: Diaspis machilicola Takahashi, 1935. Note: Syntypes, female [collected from Taiwan: Taito, Chipponsan, on Machilus sp., 23-III-1934, by R. Takahashi ( # 263-2-1–4)]. Chionaspis rotunda (Takahashi 1935) (Fig. 1D) (Takahashi 1935a; Takagi 1970; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Phenacaspis rotunda Takahashi. (1 slide) Distribution: Taiwan (Heito: Mutosan). H o s t p l a n t : Q u e rc u s p s e u d o m y r s i n i f o l i a (Fagaceae). Syn.: Phenacaspis rotunda Takahashi, 1935; Chionaspis rotunda (Takahashi 1935). Note: Holotype, female [collected from Taiwan: Takao, Heito-Gun, Mutosan, on Q u e rc u s p s e u d o m y r s in i f o l i a, 2 3 - I I I 1934, by R, Takahashi ( #39-5-3-1)]..

(3) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. 107. Table 1. A check list of the old name, current name and Chinese name of collection localities included in the paper. Old name. Current name. Chinese name. Botel Tobago. Taitung: Lanyu. 台東：蘭嶼. Chikushiko. Taipei: Zhuzihu. 台北：竹子湖. Chippon. Taitung: Zhiben (Chihpen). 台東：知本. Chipponsan. Taitung: Zhiben Mountain. 台東：知本山. Churei. Yilan: Chungling. 宜蘭：中嶺. Chushinron near Rokki. Kaohsiung: Paoshan near Liouguei. 高雄：寶山近六龜. Daichikko. Taitung: Dazhu. 台東：大竹. Garambi, Garanbi. Pingtung: Eluanbi. 屏東：鵝鑾鼻. Habon. Nantou: Holuan. 南投：合巒. Heito. Pingtung. 屏東. Hichiseizan, Hichiseisan. Taipei: Qixing Mountain (Chihsingshan). 台北：七星山. Hinode. Kaohsiung: Rizhong. 高雄：旗山：日中. Hokuto. Taipei: Beitou. 台北：北投. Hori, Horin. Nantou: Puli. 南投：埔里. Hsintien. New Taipei City: Xindian. 新北市：新店. Itabashi. New Taipei City: Banqiao. 新北市：板橋. Kaede. Taitung: Fenglin Bridge. 台東：海端：楓林橋. Kaho. Taoyuan: Hsiapu. 桃園：下埔. Kakayo, Kakayom. Taitung: Jiadai (Chiachiadai). 台東：嘉代 ( 嘉嘉代 ). Kanaron, Kanoron. Taitung: Jinlun (Chinlun). 台東：金崙. Kanshirei. Tainan: Guanziling (Kuantyling). 台南：關子嶺. Karenko. Hualien. 花蓮. Kontei. Pingtung: Kending. 屏東：墾丁. Koshun. Pingtung: Hengchun. 屏東：恆春. Kotosho. Taitung: Lanyu. 台東：蘭嶼. Kuraru. Pingtung: Sheding. 屏東：社頂. Kussha. Yilan: Kuochia. 宜蘭：庫霞. Maruyama. Taipei: Yuanshan. 台北：圓山. Murorafu, Muroruafu, Mururoafu. Yilan: Jialuo Mountain (Chialoshan). 宜蘭：加羅山. Mutosan. Pingtung: Wutoushan. 屏東：霧頭山. Oiwake. Taitung: Zhuifen (Chuifen). 台東：追分. Piyahau. Yilan: Bihou (Pihau). 宜蘭：碧候. Raisha. Pingtung: Laiyi. 屏東：來義. Riran. Taitung: Lilong. 台東：關山：里龍. Sankyo near Taihoku. New Taipei City: Sansia near Taipei. 新北市：三峽近台北. Sekinon. New Taipei City: Shimen. 新北市：石門. Sekitei. New Taipei City: Shiding (Shihding). 新北市：石碇. Shinten. New Taipei City: Xindian (Sindian). 新北市：新店. Shin-Taiheizan. Yilan: Taiping Mountain (Taipingshan). 宜蘭：太平山. Shirin. Taipei: Shilin. 台北：士林. Sozan. Taipei: Yangmingshan. 台北：陽明山.

(4) 108. 台灣農業研究第 63 卷第 2 期. Table 1. A check list of the old name, current name and Chinese name of collection localities included in the paper. (continued) Old name. Current name. Chinese name. Suisha. Nantou: Shuishe. 南投：魚池：水社. Suo. Yilan: Su’ao. 宜蘭：蘇澳. Taichu. Taichung. 台中. Taiheizan, Taihezan, Taiheisan. Yilan: Taiping Mountain (Taipingshan). 宜蘭：太平山. Taihoku, Taipei. Taipei. 台北. Tainan. Tainan. 台南. Takao Takaoshu. Kaohsiung. 高雄. Tamsui Tansui. New Taipei City: Danshui (Danshuei). 新北市：淡水. (A). (B). 50 μm. 100 μm. (C). (D). 100 μm. 100 μm. (E). (F). 100 μm. 50 μm. Fig. 1. (A) Chionaspis cinnamomicola (Takahashi 1935); (B) Chionaspis lithocarpicola Takahashi, 1942; (C) Chionaspis machilicola (Takahashi 1935); (D) Chionaspis rotunda (Takahashi 1935); (E) Chionaspis salicis (Linnaeus 1758); and (F) Chionaspis sozanica Takahashi, 1933.. Chionaspis salicis (Linnaeus 1758) (Fig. 1E) (Linnaeus 1758; Chou 1982; Ben-Dov & Miller 2013). (1) Original spelling on label: Chionaspis alni var. hokkaidensis Takah. (2 slides). Distribution: Japan (Hokkaido: Sapporo). (2) Original spelling on label: Chionaspis salicis L. (2 slides) Distribution: Turkey (Angora)..

(5) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. (3) Original spelling on label: Chionaspis salicis-niger Walsh. (2 slides) Distribution: Russia (Saghalien: Horo). Host plant: Alnus hirsuta (Betulaceae); Salix sp. (Salicaceae). Syn.: Coccus salicis Linnaeus, 1758; Coccus cryptogamus Dalman, 1826; Aspidiotes salicis (Linnaeus 1758); Aspidiotus Populi Baerensprung, 1849; Aspidiotus minimus Baerensprung, 1849; Diaspis nivea Bremi, 1849; Aspidiotus saliceti Bouché, 1851; Aspidiotus Vaccinii Bouché, 1851; Aspidiotus salicis-nigrae Walsh, 1868; Mytilaspis Maquarti Targioni Tozzetti, 1868; Mytilaspis cryptogama (Dalman 1826); Chionaspis aceris Signoret, 1869; Chionaspis alni Signoret, 1869; Chionaspis fraxini Signoret, 1869; Chionaspis vaccinii (Linnaeus 1758); Mytilaspis salicis LeBaron, 1872; Diaspis salicicorticis Bessey, 1 8 7 4 ; L e c a n i u m Va c c i n i i ( B o u c h é 1851); Aspidiotus mytilus Glaser, 1877; Coccus (Aspidiotus) mytilus (Glaser 1877); Chionaspis sorbi Douglas, 1893; Chionaspis ortholobis bruneri Cockerell, 1898; Chionaspis salicis-nigrae (Linnaeus 1758); Aspidiotus cryptogamus (Dalman 1826); Chionaspis salicis Archangelskaya, 1937; Aspidiotus aceris (Signoret 1869); Aspidiotus alni (Signoret 1869); Chionaspis minimus (Baerensprung 1849); Aspidiotus salicifex (Amyot 1847); Chionaspis salicifex (Amyot 1847); Chionaspis montana Borchsenius, 1949; Chionaspis polypora Borchsenius, 1949; Chionaspis caucasioni Hadzibejli, 1963. Chionaspis sozanica Takahashi, 1933 (Fig. 1F) (Takahashi 1933; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Chionaspis soza-. 109. nicus Takah. (3 slides) Distribution: Taiwan (Taihoku: Sozan). Host plant: Acer sp. (Aceraceae). S y n . : Tr i c h o m y t i l u s s o z a n i c a ( Ta k a h a s h i 1933); Phenacaspis sozanica (Takahashi 1933). Note: Syntypes, female [collected from Taiwan: Sozan, near Taihoku, on Acer sp., 03-VII-1932, by R. Takahashi ( # 24-3-21–3)]. Chionaspis trochodendri (Takahashi 1936) (Fig. 2A) (Takahashi 1936b; Takagi 1969; Chou 1982; Ben-Dov & Miller 2013). Original spelling on label: Phenacaspis trochodendri Takah. (4 slides) Distribution: Taiwan (Taihoku: Chikushiko, Sozan). Host plant: Trochodendron aralioides, Trochodendron sp. (Trochodendraceae). Syn.: Phenacaspis trochodendri Takahashi, 1936. Note: Syntypes, female [collected from Taiwan: Chikushiko, near Sozan, Hichiseisan, Rarasan, near Urai, on Trochodendron aralioides, 25-XII-1935, by R. Takahashi ( #40-1-1-1–40-1-1-4)]. Duplachionaspis graminis (Green 1896) (Fig. 2B) (Green 1896; Ben-Dov & Miller 2013) Original spelling on label: Chionaspis graminis Green. (8 slides) Distribution: Taiwan (Taichu; Taihoku: Shinten, Taihoku, Tansui). Host plant: Miscanthus sinensis, Poaceae, Spinifex sp. (Poaceae). Syn.: Chionaspis graminis Green, 1896; Trichomytilus graminis (Green 1896); Poliaspis graminis (Green 1896). Duplachionaspis natalensis (Maskell 1896) (Fig. 2C) (Tao 1989; Ben-Dov & Miller 2013).

(6) 110. 台灣農業研究第 63 卷第 2 期. (A). (B). 100 μm. 100 μm. (D). (C). 100 μm. 50 μm. (E). (F). 100 μm. 100 μm. Fig. 2. (A) Chionaspis trochodendri (Takahashi 1936); (B) Duplachionaspis graminis (Green 1896); (C) Duplachionaspis natalensis (Maskell 1896); (D) Greenaspis bambusifoliae (Takahashi 1930); (E) Greenaspis elongata (Green 1896); and (F) Kuwanaspis annandalei (Green 1919).. Original spelling on label: Chionaspis stanotophri Cooley. (5 slides) Distribution: Taiwan (Taihoku; Tainan; Taito: Kanoron). Host plant: Phragmites sp., Poaceae, Saccharum officinarum (Poaceae). Syn.: Chionaspis spartinae natalensis Maskell, 1896; Chionaspis stanotophri Cooley, 1899; Chionaspis natalensis Maskell, 1896; Chionaspis (Phenacaspis) natalensis (Maskell 1896); Chionaspis graminis aegyptiaca Hall, 1923; Trichomytilus stanotophri (Cooley 1899); Polyaspis stanotophri (Cooley 1899); Poliaspis stanotophri (Cooley 1899); Duplachionaspis stanotophri (Cooley. 1899); Duplachionaspis herbicola Mamet, 1959. Greenaspis bambusifoliae (Takahashi 1930) (Fig. 2D) (Takahashi 1930; Tao 1989; Wong et al. 1999; Ben-Dov & Miller 2013). Original spelling on label: Chionaspis bambusifoliae Takahashi. (11 slides) Distribution: Taiwan (Nantou: Hori, Suisha; Taihoku). Host plant: Bambusa sp. (Poaceae). Syn.: Chionaspis bambusifoliae Takahashi, 1930. Note: Syntypes, female [collected from Taiwan: Taihoku, on Bambusa sp., I-1930, by R.Takahashi ( # 23-2-2-1–2、 # 23-2-24–5)]..

(7) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. Greenaspis elongata (Green 1896) (Fig. 2E) (Green 1896; Tao 1989; Wong et al. 1999; Ben-Dov & Miller 2013). (1) Original spelling on label: Chionaspis elongata (Green 1896) (15 slides) Distribution: Taiwan (Taihoku: Hichiseizan, Hokuto, Itabashi, Maruyama, Sozan, Taihoku; Taito: Riran). (2) Original spelling on label: Greenaspis elongata Green, 1896 (1 slide) Distribution: Malaysia (Malaya: Penang). Host plant: Bambusa sp. (Poaceae). Syn.: Mytilaspis elongata Green, 1896; Chionaspis elongata (Green 1896); Tr i c h o m y t i l u s e l o n g a t u s ( G r e e n 1896); Greenaspis yunnanensis Ferris, 1952.. (A). 111. Kuwanaspis annandalei (Green 1919) (Fig. 2F) (Green 1919; Ben-Dov & Miller 2013) Original spelling on label: Chionaspis annandalei Green. (1 slide) Distribution: Taiwan (Giran: Suo). Host plant: Poaceae. Syn.: Chionaspis annandalei Green, 1919. Kuwanaspis arundinariae Takahashi, 1938 (Fig. 3A) (Takahashi 1938; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Kuwanaspis arundinariae Takah. (4 slides) Distribution: Taiwan (Giran: Taiheizan). Host plant: Arundinaria sp. (Poaceae). Syn.: Nikkoaspis arundinariae (Takahashi 1938). Note: Syntypes, female [collected from Tai-. (B). 100 μm. (C). (D). 100 μm. 100 μm. (E). 100 μm. 50 μm. (F). 100 μm. Fig. 3. (A) Kuwanaspis arundinariae Takahashi, 1938; (B) Kuwanaspis bambusifoliae (Takahashi 1934); (C) Kuwanaspis elongata (Takahashi 1930); (D) Kuwanaspis linearis (Green 1922); (E) Kuwanaspis neolinearis (Takahashi 1930); and (F) Kuwanaspis phragmitis (Takahashi 1931)..

(8) 112. 台灣農業研究第 63 卷第 2 期. wan: Taiheizan, on Arundinaria sp., 14X-1937 by R. Takahashi ( #31-1-1-1–4)]. Kuwanaspis bambusifoliae (Takahashi 1934) (Fig. 3B) (Takahashi 1934; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Tsukushiaspis bambusifoliae Takah. (2 slides) Distribution: Taiwan (Taihoku: Hichiseizan). Host plant: Bambusa sp. (Poaceae). Syn.: Tsukushiaspis bambusifoliae Takahashi, 1934. Note: Syntypes, female [collected from Taiwan: Taihoku, on Mt. Hichisei, on Bambusa sp., 27-III-1932 and 23-XI-1933, by R. Takahashi ( #31-1-2-4–5)]. Kuwanaspis elongata (Takahashi 1930) (Fig. 3C) (Takahashi 1930; Tao 1989; Wong et al. 1999; Ben-Dov & Miller 2013). Original spelling on label: Tsukushiaspis elongata Takahashi. (6 slides) Distribution: Taiwan (Nantou: Suisha; Taihoku: Maruyama). Host plant: Bambusa sp., Dendrocalamus sp. (Poaceae). Syn.: Tsukushiaspis elongata Takahashi, 1930. Note: Syntypes, female [collected from Taiwan: Suisha, on Bambusa sp., 2-X-1929, by R. Takahashi ( #31-1-3-1–5)]. Kuwanaspis linearis (Green 1922) (Fig. 3D) (Takahashi 1942a; Chou 1982; Ben-Dov & Miller 2013). Original spelling on label: Kuwanaspis linearis Green. (3 slides) Distribution: Malaysia (Malaya: Kuala Lumpur); Singopore. Host plant: Bambusa sp. (Poaceae). Syn.: Chionaspis linearis Green, 1922; Tsukushiaspis linearis (Green 1922). Kuwanaspis neolinearis (Takahashi 1930) (Fig. 3E) (Takahashi 1930, 1942a; Tao & Wong 1982; Tao 1989; Wong et al. 1999; Ben-Dov & Miller 2013). (1) Original spelling on label: Chionaspis. neolinearis Takah. (2 slides) Distribution: Taiwan (Nantou: Suisha). Note: Syntype, female [collected from Taiwan: Taihoku, on Bambusa sp., 02-X1929, by R. Takahashi ( #23-5-1-2)]. (2) Original spelling on label: Tsukushiaspis neolinearis Takah. (12 slides) Distribution: Singapore; Taiwan (Giran: Piyahau; Heito: Heito, Kuraru; Hualien: Karenko; Taihoku). Host plant: Bambusa sp. (Poaceae). Syn.: Tsukushiaspis neolinearis Takahashi, 1930; Chuaspis neolinearis (Takahashi 1930). Kuwanaspis phragmitis (Takahashi 1931) (Fig. 3F) (Takahashi 1931; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Tsukushiaspis phragmitis Takahashi. (6 slides) Distribution: Taiwan (Giran: Suo; Taihoku: S e k i n o n , Ta m s u i ; Ta i t o : C h i p p o n , Daichikko) Host plant: Artemisia feddei (Compositae); Phragmites sp. (Poaceae). Syn.: Tsukushiaspis phragmitis Takahashi, 1931. Note: Syntypes, female [collected from Taiwan: Suo, Chippon, Daichikko near Daibu, on Phragmites sp., 30-XII-1924, by R. Takahashi ( #31-3-4-1 –2)]. Kuwanaspis pseudoleucaspis (Kuwana 1902) (Fig. 4A) (Kuwana 1902; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Tsukushiaspis bambusae Kuwana. (2 slides) Distribution: Taiwan (Heito: Raisha). Host plant: Bambusa sp. (Poaceae). Syn.: Leucaspis bambusae Kuwana, 1902; Lepidosaphes bambusae (Kuwana 1902); Mytilaspis bambusae (Kuwana 1902); Chionaspis pseudoleucaspis (Kuwana 1902); Tsukushiaspis bam-.

(9) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. (A). 113. (B). 100 μm 100 μm. (C). (D). 50 μm. (E). 100 μm. (F). 100 μm. 200 μm. Fig. 4. (A) Kuwanaspis pseudoleucaspis (Kuwana 1902); (B) Kuwanaspis suishana (Takahashi 1930); (C) Kuwanaspis vermiformis (Takahashi 1931); (D) Ledaspis atalantiae (Takahashi 1933); (E) Neoquernaspis lithocarpi (Takahashi 1931); and (F) Nikkoaspis formosana (Takahashi 1930).. busae (Kuwana 1902); Tsukushiaspis pseudoleucaspis (Kuwana 1902); Lepidosaphoides bambusae (Kuwana 1902); Kuwanaspis bambusae (Kuwana 1902). Kuwanaspis suishana (Takahashi 1930) (Fig. 4B) (Takahashi 1930, 1942b; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Tsukushiaspis shuishanus Tak. (15 slides) Distribution: Taiwan (Hualien: Karenko; Nantou: Suisha; Taihoku: Maruyama, Taihoku); Thailand (Chingmai). Host plant: Bambusa sp., Bambusa stenostachya. (Poaceae). Syn.: Tsukushiaspis suishanus Takahashi, 1930; Kuwanaspis suishanus (Takahashi 1930).. Kuwanaspis vermiformis (Takahashi 1931) (Fig. 4C) (Takahashi 1931, Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Tsukushiaspis vermiformis Takah. (4 slides) Distribution: Taiwan (Taihoku: Maruyama, Taihoku). Host plant: Bambusa sp., Bambusa stenos t a c h y a , D e n d ro c a l a m u s l a t i f l o r u s . (Poaceae). Syn.: Tsukushiaspis vermiformis Takahashi, 1931. Note: Syntypes, female [collected from Taiwan: Taihoku, Tosei, on Bambusa stenostachya, Bambusa sp., Dendrocalamus latiflorus ( #31-5-3-1–3)]..

(10) 114. 台灣農業研究第 63 卷第 2 期. Ledaspis atalantiae (Takahashi 1933) (Fig. 4D) (Takahashi 1933; Ben-Dov & Miller 2013) (1) Original spelling on label: Chionaspis atalantiae Takah. (7 slides) D i s t r i b u t i o n : Ta i w a n ( H e i t o : K o n t e i ; Takao). Note: Syntypes, female [collected from Ta i w a n : K o n t e i , n e a r K o s h u n , o n Atalantia lilocularis, 24-V-1932, by R.Takahashi ( #23-1-2-1–23-1-3-1)]. (2) Original spelling on label: Dinaspis atalantiae Takah. (4 slides) Distribution: Taiwan (Heito: Garanbi). Host plant: Atalantia lilocularis, Severinia buxifolia (Rutaceae). Syn.: Chionaspis atalantiae Takahashi, 1933; Trichomytilus atalantiae (Takahashi 1933); Phenacaspis atalantiae (Takahashi 1933); Pseudaulacaspis atalantiae (Takahashi 1933).. hashi 1930). Note: Syntypes, female [collected from Taiwan: Suisha, on Bambusa sp., 23-XI1929 ( # 31-2-1-1–2) and 26-I-1930 ( # 312-1-4–5) by R. Takahashi]. Nikkoaspis hichiseisana (Takahashi 1934) (Fig. 5A) (Takahashi 1934; Takagi 1961; Tao 1989; Ben-Dov & Miller 2013). Original spelling on label: Tsukushiaspis hichiseisana Takah. (8 slides) Distribution: Taiwan (Taihoku: Hichiseizan). Host plant: Bambusa sp. (Poaceae). Syn.: Tsukushiaspis hichiseisana Takahashi, 1934; Kuwanaspis hichiseisana (Takahashi, 1934). Note: Syntypes, female [collected from Taiwan: Mt. Hichisei, near Taihoku, on Bambusa sp., 23-IX-1933 by R. Takahashi ( #31-2-2-1–31-2-3-3)]. Nikkoaspis sasae (Takahashi 1936) (Fig. 5B). Neoquernaspis lithocarpi (Takahashi 1931) (Fig. 4E) (Takahashi 1931; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Pinnaspis lithocarpi Takah. (6 slides) Distribution: Taiwan (Nantou: Hori, Suisha). Host plant: Lithocarpus sp., Pasania ternaticupula. (Fagaceae) Syn.: Pinnaspis lithocarpi Takahashi, 1931; Quernaspis lithocarpi (Takahashi 1931). Note: Syntypes, female [collected from Taiwan: Suisha, on Lithocarpus sp., 23-XI-1929, by R. Takahashi ( #41-3-3-1–2)].. Original spelling on label: Tsukushiaspis sasae Tak. (3 slides) TARI specimens Distribution: China (Zhejiang: Hangzhou, Huchow). Host plant: Bambusa sp., Sasa sp. (Poaceae). Syn.: Tsukushiaspis sasae Takahashi, 1936; Kuwanaspis sasae (Takahashi 1936). Note: Syntypes, female [collected from China: Zhejiang, Huchow, on Sasa sp., VIII1935, by S.P. Chu ( #31-4-2-1–2)].. Nikkoaspis formosana (Takahashi 1930) (Fig. 4F) (Takahashi 1930, 1934; Tao 1989; Ben-Dov & Miller 2013) Original spelling on label: Tsukushiaspis formosanus Takah. (5 slides) Distribution: Taiwan (Nantou: Suisha). Host plant: Bambusa sp. (Poaceae). Syn.: Tsukushiaspis formosanus Takahashi, 1930; Kuwanaspis formosana (Taka-. Nikkoaspis shiranensis Kuwana, 1928 (Fig. 5C) (Takahashi 1934; Ben-Dov & Miller 2013) Original spelling on label: Kuwanaspis shiranensis Kuwana. (1 slide) Distribution: Japan (Hokkaido: Sapporo). Host plant: Sasa veitchii (Poaceae). Syn.: Tsukushiaspis shiranensis (Kuwana 1928); Kuwanaspis shiranensis (Kuwana 1928).. (Takahashi 1936a; Takagi 1961; Chou 1982; Ben-Dov & Miller 2013).

(11) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. (A). 115. (B). 100 μm 100 μm. (D). (C). 50 μm. 200 μm. (E). (F). 100 μm. 50 μm. Fig. 5. (A) Nikkoaspis hichiseisana (Takahashi 1934); (B) Nikkoaspis sasae (Takahashi 1936); (C) Nikkoaspis shiranensis Kuwana, 1928; (D) Pinnaspis aspidistrae aspidistrae (Signoret 1869); (E) Pinnaspis javanensis (Kuwana in Kuwana & Muramatsu 1931); and (F) Pinnaspis siamensis Takahashi, 1942.. Pinnaspis aspidistrae aspidistrae (Signoret 1869) (Fig. 5D) (Lindinger 1912; Tao 1989; Wong et al. 1999; Ben-Dov & Miller 2013) (1) Original spelling on label: Hemi-Chionaspis aspidistrae Signt. (1 slide) Distribution: Taiwan (Taihoku). (2) Original spelling on label: Pinnaspis aspidistrae Signt. (44 slides) Distribution: China (Foochow); Fiji; Japan (Kyushu: Kagoshima); Okinawa; Malaysia (Malaya: Bukit Asahan, G. Batu Brinchang, Kuala Lumpur, Serdang, Taiping); Taiwan (Giran: Churei; Heito: Garanbi; Hualien: Karenko; Nantou: Habon Near Urai, Suisha; Taihoku: Hinokiyama, Shirin, Taihoku; Tainan; Taito: Kaede,. Kakayo, Oiwake; Takaoshu: Hinode); Thailand (Ubon). Host plant: Asplenium sp. (Aspleniaceae); Citrus sp. (Rutaceae); Cyathea sp., Cycas sp. (Cycadaceae); Cymbidium ensifolium, Orchid (Orchidaceae), Daemonorops sp., Elaeis guineensis (Arecaceae); Diospyros sp. (Ebenaceae); Elatostema sp. (Urticaceae); Fern sp. (Psilotaceae); Ficus sp., Ficus pumila, Ficus retusa, Ficus tinctoria (Moraceae); Hydrangea sp. (Saxifragaceae); Pandanus sp., Pandanus atrocarpus (Pandanaceae); Podocarpus sp. (Podocarpaceae), Rhododendron probably jasminiflorum (Ericaceae); Smilax sp. (Smi-.

(12) 116. 台灣農業研究第 63 卷第 2 期. lacaceae); Styrax sp. (Styracaceae); Viburnum sp. (Caprifoliaceae). Syn.: Chionaspis aspidistrae Signoret, 1869; Chionaspis brasiliensis Signoret, 1869; Chionaspis latus Cockerell, 1896; Hemichionaspis aspidistrae brasiliensis (Signoret 1869); Hemichionaspis aspidistrae lata (Signoret 1869); Chionaspis (Pinnaspis) aspidistrae (Signoret 1869); Pinnaspis (Hemichionaspis) aspidistrae (Signoret 1869); Pinnaspis brasiliensis (Signoret 1869); Pinnaspis ophiopogonis Takahashi, 1952; Pinnaspis caricis Ferris, 1957.. Pinnaspis siamensis Takahashi, 1942 (Fig. 5F) (Takahashi 1942b; Ben-Dov & Miller 2013) Original spelling on label: Pinnaspis lithocarpi var. siamensis Tak. (8 slides) Distribution: Thailand (Mt. Sutep; Siam). Syn.: Pinnaspis lithocarpi siamensis Takahashi, 1942; Quernaspis siamensis (Takahashi 1942). Host plant: Lithocarpus sp. (Fagaceae) Note: Syntypes, female [collected from Thailand: Mt. Sutep, on Lithocarpus sp. 09I V- 1 9 4 0 , b y R . Ta k a h a s h i ( # 4 1 - 4 - 1 1–41-4-2-3)].. pis minor Mask. (1 slide) Distribution: Taiwan (Taihoku: Taihoku). (2) Original spelling on label: Pinnaspis minor Mask. (12 slides) Distribution: French Polynesia (Tahiti); Micronesia (Caroline Isls.: Ponape: Kolonia); Taiwan (Heito: Koshun; Ta i h o k u : H i c h i s e i z a n ; Ta o y u a n : Kaho); Thailand (Bangkok). Host plant: Chelidonium sp. (Papaveraceae); Cocos nucifera (Arecaceae); Eurya sp. (Theaceae); Planchonella sp. (Sapotaceae); Solanum sp. (Solanaceae). Syn.: Hemichionaspis minor strachani C o o l e y, 1 8 9 9 ; H e m i c h i o n a s p i s Marchali Cockerell, 1902; Hemichionaspis townsendi Cockerell, 1905; Chionaspis (Hemichionaspis) aspidistrae gossypii Newstead, 1908; Hemichionaspis aspidistrae gossypii (Newstead 1906); Hemichionaspis proxima Leonardi, 1914; Hemichionaspis (Pinnaspis) marchali (Cockerell 1902); Pinnaspis minor (Cooley 1899); Pinnaspis minor strachani (Cooley 1899); Pinnaspis proxima (Leonardi 1914); Pinnaspis (Hemichionaspis) aspidistrae gossypii (Newstead 1908); Pinnaspis temporaria Ferris, 1942; Pinnaspis aspidistrae gossypii (Newstead 1908); Pinnaspis gossypii (Newstead 1908); Pinnaspis marchali (Cockerell 1902); Hemichionaspis gossypii (Newstead 1908); Chionaspis (Hemichionaspis) gossypii (Newstead 1908); Pinnaspis townsendi (Cockerell 1905).. Pinnaspis strachani (Cooley 1899) (Fig. 6A). Pinnaspis theae (Maskell 1891) (Fig. 6B) (Taka-. Pinnaspis javanensis (Kuwana 1931) (Fig. 5E) (Takagi 1985; Ben-Dov & Miller 2013) Original spelling on label: Pinnaspis javanensis Kuwana. (1 slide) Distribution: Malaysia (Malaya: Sunge Buloh). Host plant: Hevea sp. (Euphorbiaceae). Syn.: Chionaspis javanensis Kuwana, 1931.. (Tao 1989; Ben-Dov & Miller 2013). hashi 1929; Tao 1989; Ben-Dov & Miller 2013). (1) Original spelling on label: Hemichionas-. Original spelling on label: Pinnaspis theae.

(13) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. (A). 117. (B). 100 μm 50 μm. (C). (D). 100 μm. 50 μm. (E). (F). 100 μm. 100 μm. Fig. 6. (A) Pinnaspis strachani (Cooley 1899); (B) Pinnaspis theae (Maskell 1891); (C) Pinnaspis uniloba (Kuwana 1909); (D) Unaspis acuminata (Green 1896); (E) Unaspis citri (Comstock 1883); and (F) Unaspis euconymi (Comstock 1881).. Mask. (3 slides) Distribution: Taiwan (Heito: Koshun; Tainan: Kanshirei; Takao). Host plant: Cleyera japonica, Thea japonica, Thea sp. (Theaceae). Syn.: Chionaspis theae Maskell, 1891; Aspidiotus theae (Maskell 1891); Chionaspis aspidistrae theae (Maskell 1891); Hemichionaspis theae (Maskell 1891); Chionaspis separata Green, 1900; Hemichionaspis separata (Green 1900); Trichomytilus theae (Maskell 1891); Pinnaspis separata (Green 1900). Pinnaspis uniloba (Kuwana 1909) (Fig. 6C). (1) Original spelling on label: Pinnaspis simplex Ferris. (3 slides) Distribution: Taiwan (Tainan: Kanshirei, Sekitei). (2) Original spelling on label: Pinnaspis uniloba Kuwana. (1 slide) Distribution: Taiwan (Taihoku: Sozan). Host plant: Camellia sp., Cleyera japonica (Theaceae). Syn.: Mytilaspis (Lepidosaphes) uniloba Kuwana, 1909; Lepidosaphes uniloba (Kuwana 1909); Jaapia uniloba (Kuwana 1909); Lepidaspidis uniloba (Kuwana 1909); Pinnaspis simplex Ferris, 1921.. (Ferris 1921; Takahashi 1929; Tao 1989; Wong et al. 1999; Ben-Dov & Miller 2013). Unaspis acuminata (Green 1896) (Fig. 6D).

(14) 118. 台灣農業研究第 63 卷第 2 期. (Green & Laing 1923); Diaspis annae (Malenotti 1916); Chionaspis annae (Malenotti 1916).. (Green 1896; Chou 1982; Ben-Dov & Miller 2013). Original spelling on label: Unaspis acuminata Green. (2 slides) Distribution: Thailand (Chiengsen; Mt. Sutep). Host plant: Mangifera indica (Anacardiaceae). Syn.: Chionaspis acuminata Green, 1896. Unaspis citri (Comstock 1883) (Fig. 6E) (Chou 1982; Ben-Dov & Miller 2013). (1) Original spelling on label: Dinaspis veitchi Green et Laing. (1 slide) Distribution: Papua New Guinea (Admiralty Isls.: Manus). (2) Original spelling on label: Prontaspis citri Comst. (1 slide) Distribution: Malaysia (Malaya). Host plant: Citrus sp. (Rutaceae). Syn.: Chionaspis citri Comstock, 1883; H o w a rd i a c i t r i ( C o m s t o c k 1 8 8 3 ) ; Chionaspis citricola (Comstock 1883); Dinaspis annae Malenotti, 1 9 1 6 ; P ro n t a s p i s c i t r i ( C o m s t o c k 1883); Dinaspis veitchi Green & Laing, 1923; Trichomytilus veitchi. (A). Unaspis euconymi (Comstock 1881) (Fig. 6F) (Chou 1982; Ben-Dov & Miller 2013). Original spelling on label: Unaspis nakayamai Takah. et Kanda. (1 slide) Distribution: Korea (Suigen). Host plant: Ilex serrata (Aquifoliaceae). Syn.: Chionaspis euonymi Comstock, 1881; Unaspis euonymi (Comstock 1881); Unaspis nakayamai Takahashi & Kanda, 1939. Unaspis kanoi (Takahashi 1935) (Fig. 7A) (Takahashi 1935b; Ben-Dov & Miller 2013). Original spelling on label: Lepidosephas (Coccomytilus) kanoi Takah. (5 slides) Distribution: Taiwan (Taito: Kotosho). Syn.: Coccomytilus kanoi Takahashi, 1935. Note: Syntypes, female [collected from Taiwan: Botel Tobago, on undetermined tree, 1935, by T. Kano ( #33-3-4-1–5)].. (B). 100 μm. 50 μm. (C). 200 μm. Fig. 7. (A) Unaspis kanoi (Takahashi 1935); (B) Unaspis turpiniae Takahashi, 1934; and (C) Unaspis yanonensis (Kuwana 1923)..

(15) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. Unaspis turpiniae Takahashi, 1934 (Fig. 7B) (Takahashi 1934; Tao 1989; Ben-Dov & Miller 2013). Original spelling on label: Dinaspis acuminata var. turpiniae Takah. (2 slides) Distribution: Taiwan (Giran: Muroruafu). Host plant: Ilex sp. (Aquifoliaceae). Syn.: Unaspis acuminata turpiniae Takahashi, 1934; Chionaspis turpinae (Takahashi 1934). Unaspis yanonensis (Kuwana 1923) (Fig. 7C) (Takahashi & Kanda, 1939; Chou 1982; Ben-Dov & Miller 2013). Original spelling on label: Prontaspis yanonensis Kuw. (2 slides) Distribution: Taiwan (Taihoku: Sankyo near Taihoku). Host plant: Citrus sp. (Rutaceae). Syn.: Prontaspis yanonensis Kuwana, 1923.. DISCUSSION Based on the modern taxonomic system, the number of Chionaspinae species collected and deposited at TARI Insect Collection is 39. The present paper could be a database to set up the online information of scientific name, synonyms, host plants, distribution and digital archives through Digital Insect of TARI website (http://digiins.tari.gov.tw/) which will meet the requirement of both basic and applied works and will be helpful to taxonomic researchers, quarantine inspectors and farmers in the future.. ACKNOWLEDGMENTS We would like to thank the National Science Council (grant numbers 99-2631-H-055001 and 100-2631-H-055-001) and Council of Agriculture, Executive Yuan (grant number 102AS-7.1.1-CI-C1) for financial support.. REFERENCES Ben-Dov, Y. and D. Miller. 2013. ScaleNet: A database of. 119. the scale insects of the world. (http://www.sel.barc. usda.gov/scalenet/scalenet.htm). Borchsenius, N. S. 1966. A Catalogue of the Armoured Scale Insects (Diaspidoidea) of the World. Nauka. Moscow & Leningrad. 449 pp. (in Russian) Chou, I. 1982. Monograph of the Diaspididae of China. Vol. 1. Shanxi Publishing House of Science and Technology. Shanxi. 195 pp. (in Chinese) Chu, Y. I. and T. Yamanaka. 1973. A check list of the present and old name of insect collect localities in Taiwan. J. Taiwan Mus. 16:31–72. Chu, Y. I. and T. Yamanaka. 1974. A check list of the present and old name of insect collect localities in Taiwan (continued). J. Taiwan Mus. 17:51–120. Chu, Y. I. and T. Yamanaka. 1975. A check list of the present and old name of insect collect localities in Taiwan (continued). J. Taiwan Mus. 17:121–150. Ferris, G. F. 1921. Some Coccidae from Eastern Asia. Bull. Entomol. Res. 12:211–220. Green, E. E. 1896. Catalogue of Coccidae collected in Ceylon. Indian Mus. Notes 4:2–10. Green, E. E. 1919. Notes on Indian Coccidae of the subfamily Diaspidinae, with description of new species. Records Indian Mus. 16:433–449. Kuwana, S. I. 1902. Coccidae (scale insects) of Japan. Proc. Calif. Acad. Sci. 3:43–98. Lindinger, L. 1912. Die Schildläuse (Coccidae) Europas, Nordafrikas und Vorder-Asiens, einschliesslich der Azoren, der Kanaren und Madeiras. Ulmer. Stuttgart. 388 pp. (in German) Linnaeus, C. 1758. Insecta Hemiptera–Coccus. Systema naturae. Salvii Holmiae. Stockholm. 823 pp. Takagi, S. 1961. A contribution to the knowledge of the Diaspididini of Japan (Homoptera: Coccoidea). Part II. Insecta Matsumurana 24:4–42. Takagi, S. 1969. Diaspididae of Taiwan based on material collected in connection with the Japan-U.S. Cooperative science programme, 1965 (Homoptera: Coccoidea). Part I. Insecta Matsumurana 32:1–110. Takagi, S. 1970. Diaspididae of Taiwan based on material collected in connection with the Japan-U.S. Cooperative Science Programme, 1965 (Homoptera: Coccoidea). Part II. Insecta Matsumurana 33:1–142. Takagi, S. 1985. The scale insect genus Chionaspis; A revised concept (Homoptera: Coccoidea: Diaspididae). Insecta Matsumurana (n.s.) 33:1–77. Takahashi, R. 1929. Observations on the Coccidae of Formosa. Part I. Formosa Govt. Res. Inst. Dept. Agric. Rept. 40. Taihoku. 80 pp..

(16) 120. 台灣農業研究第 63 卷第 2 期. Takahashi, R. 1930. Observations on the Coccidae of Formosa. Part II. Formosa Govt. Res. Inst. Dept. Agric. Rept. 43. Taihoku. 45 pp.. Takahashi, R. 1942a. Some Coccidae from Malaya and Hongkong (Homoptera). Trans. Nat. Hist. Soc. Formosa 32:63–68.. Takahashi, R. 1931. Descriptions of some new Formosan Coccidae. (Rhynchota). Bull. Entomol. Res. 22:211–220.. Takahashi, R. 1942b. Some injurious insects of agricultural plants and forest trees in Thailand and IndoChina. Part II. Coccidae Report. Formosa Govt. Res. Inst. Dept. Agric. Rept. 81. Taihoku. 56 pp.. Takahashi, R. 1933. Observations on the Coccidae of Formosa. Part III. Formosa Govt. Res. Inst. Dept. Agric. Rept. 60. Taihoku. 64 pp. Takahashi, R. 1934. Observations on the Coccidae of Formosa, Part IV. Formosa Govt. Res. Inst. Dept. Agric. Rept. 63. Taihoku. 38 pp. Takahashi, R. 1935a. Observations on the Coccidae of Formosa. Part V. Formosa Govt. Res. Inst. Dept. Agric. Rept. 66 .Taihoku. 37 pp. Takahashi, R. 1935b. Scale insects of the genus Coccomytilus from Formosa. Trans. Nat. Hist. Soc. Formosa 25:444–447. Takahashi, R. 1936a. Some Coccidae from China (Hemiptera). Peking Nat. Hist. Bull. 10:217–222. Takahashi, R. 1936b. Some Coccidae from Formosa and Japan (Homoptera). Part I. Mushi 9:1–8. Takahashi, R. 1938. Some Coccidae from Formosa and Japan (Homoptera). Part III. Mushi 11:42–45.. Takahashi, R. and S. Kanda. 1939. A new species of Coccidae (Homoptera) from Korea. Annot. Zool. Jpn. 18:185–187. Tao, C. C. C. 1978. Check list and host plant Index to scale insects of Taiwan, Republic of China. J. Agric. Res. China 27:77–141. (in Chinese with English abstract) Tao, C. C. C. 1989. Scale insects name list of Taiwan, Republic of China. Bull. Taichung Dist. Agric. Improve. Stat. 22:57–70. (in Chinese with English abstract) Tao, C. C. C. and C. Y. Wong. 1982. Description of a new genus and species of bamboo scale Chuaspis shuichuensis (Homoptera: Coccoidea: Diaspididae) from Taiwan, China. Qtrly J. Taiwan Mus. 35:123–125. Wong, C. Y., S. P. Chen, and L. Y. Chou. 1999. Guidebook to Scale Insects of Taiwan. Agric. Res. Inst. Pub. No. 89. Taichung. 98 pp. (in Chinese).

(17) List of Chionaspinae (Hemiptera: Sternorrhyncha: Coccoidea: Diaspididae) in TARI. 121. 農業試驗所昆蟲標本館雪盾介殼蟲亞科 (半翅目：胸喙亞目：介殼蟲總科：盾介殼蟲科) 標本名錄陳淑佩 1,* 翁振宇 2 吳文哲 3,4 摘要陳淑佩、翁振宇、吳文哲。2014。農業試驗所昆蟲標本館雪盾介殼蟲亞科 (半翅目：胸喙亞目：介殼蟲總科：盾介殼蟲科) 標本名錄。台灣農業研究 63(2):105–121。本文羅列 9 屬 39 種 (246 片玻片標本) 農業試驗所館藏之雪盾介殼蟲亞科 [半翅目 (Hemiptera) 胸喙亞目 (Sternorrhyncha) 介殼蟲總科 (Coccoidea) 盾介殼蟲科 (Diaspididae)] 名錄，其中亦包含異名、寄主植物、採集分佈及標本形態圖等資料。關鍵詞：盾介殼蟲、農業試驗所、館藏標本。. * 1 2 3 4. 投稿日期：2014 年 1 月 24 日；接受日期：2014 年 3 月 26 日。通訊作者：spchen@tari.gov.tw 農委會農業試驗所應用動物組副研究員。台灣台中市。農委會農業試驗所應用動物組前技工。台灣台中市。國立台灣大學昆蟲學系教授。台灣台北市。國立台灣大學生物資源暨農學院植物醫學研究中心主任。台灣台北市。.

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